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Darwin proudly references his son George’s technical drawing.

Darwin publicly thanks Lady Dorothy Nevill for providing orchid samples.

Descent is Darwin’s second book on evolutionary theory. In Descent Darwin applies the evolutionary theory of On the Origin of the Species (1859) to human evolution; he provides detailed evidence of the relationship between man and other forms and…

Lucy Wedgwood’s work on Oxlips was acknowledged in Darwin’s Different Forms of Flowers but she was identified only as “a friend in Surrey”.

In March 1871, Darwin also contacted the German-born painter and illustrator, Josef Wolf (1820–99), a specialist on animals who had arrived in London in 1848 and had already illustrated many natural history publications. Darwin had asked the Regent’s…

Like Descent, the main consideration of Expression is human origins. In Expression Darwin focused on human emotion and psychology, exploring the origin of genetically determined aspects of human behaviour and emotion.

Darwin publicly acknowledges his son in law Richard Litchfield’s contribution to Expression on the subject of music.

Darwin met the Swedish photographer Oscar Rejlander, famous for his character studies, in 1871. Rejlander, who was based in London, went on to provide twenty of the images published in Expression, more than half the photographs in the book.

I have not been able to catch her in a reflecting mood, to make yr observation but she told me a fact which I think quite worthy to go down in your book along with the baby’s nods & winks viz. that when she coughs very sharply in the dark sparks come…

Like all of Darwin’s children, Francis helped his father from a young age. As a child he and his siblings made observations and collected plant samples. Later in life Francis, classically-educated in Latin, proved a useful source of help for Darwin’s…