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Darwin met the Swedish photographer Oscar Rejlander, famous for his character studies, in 1871. Rejlander, who was based in London, went on to provide twenty of the images published in Expression, more than half the photographs in the book.

Darwin publicly acknowledges his son in law Richard Litchfield’s contribution to Expression on the subject of music.

Like Descent, the main consideration of Expression is human origins. In Expression Darwin focused on human emotion and psychology, exploring the origin of genetically determined aspects of human behaviour and emotion.

In March 1871, Darwin also contacted the German-born painter and illustrator, Josef Wolf (1820–99), a specialist on animals who had arrived in London in 1848 and had already illustrated many natural history publications. Darwin had asked the Regent’s…

Lucy Wedgwood’s work on Oxlips was acknowledged in Darwin’s Different Forms of Flowers but she was identified only as “a friend in Surrey”.

Descent is Darwin’s second book on evolutionary theory. In Descent Darwin applies the evolutionary theory of On the Origin of the Species (1859) to human evolution; he provides detailed evidence of the relationship between man and other forms and…

Darwin publicly thanks Lady Dorothy Nevill for providing orchid samples.

Darwin proudly references his son George’s technical drawing.

Darwin refers to Mary Treat’s published work ‘Observations on the Sundew’, American Naturalist Vol. VII, (December, 1873), pp. 705 – 708.

Darwin praises the work of Mary Treat, an entomologist from New Jersey. Her article on Drosera was referenced by Darwin in his 1875 publication Insectivorous Plants. As a published naturalist, Treat had already constructed a public profile for…